Studies in Mathematics, University of Oxford; PhD in Biomathematics, Imperial College, London. Formerly: with National Institute for Medical Research; 1990, Royal Society University Research Fellow, University of Oxford; 1994, secondment to Institut Pasteur, Paris; 1998, Head, Mathematical Biology, BBSRC's Institute for Animal Health; 2000, University of Personal profile Oxford; 2004, Professor of Mathematical Biology, University of Oxford; since 2005, current position. 2008, Senior Research Fellowship, All Souls College, University of Oxford. Member: Science Advisory Council, Department for the Environment and Rural Affairs; National Expert Panel for New and Emerging Infections, Department of Health. Research interests: use of mathematical models to aid understanding of evolution and spread of infectious agents.