PhD, Oxford University; postdoctoral work in Heidelberg, Paris and at Bell Labs. Since 1997, at UCL; 2001, Professor of Neuroscience; supported by the Wellcome Trust (Career Development Fellowship 1997; Senior Research Fellowship 2001, 2006; Principal Research Fellowship 2011) and the European Research Council (Advanced Grant 2010). Member, EMBO. Fellow: Academy of Medical Sciences; Royal Society.